The Starfleet Corps of Engineers crew on the U.S.S. da Vinci faces their biggest challenge yet: moving on after a tragedy. With new personnel on board and Commander Sonya Gomez still grieving over the loss of Kieran Duffy, the S.C.E. tackles new missions that take them from the streets of San Francisco to the Venus Terraforming Project, from Ferenginar's past to the depths of a black hole, and from the dark days of the Dominion War to a present-day encounter with the Androssi.